Stardate not given: Jadzia Dax, who always thought she was musically untalented, plays a melody on a keyboard that she seems to remember from somewhere and begins to hallucinate. Dr. Bashir examines her and finds dangerously low levels of a neurotransmitter between the Jadzia host and the Dax symbiont. They take the Defiant to the Trill homeworld to consult Dr. Renhol of the Symbiosis Commission. Timor, one of the unjoined Trills taking care of the symbionts senses there is something wrong with one of the Dax hosts. Dax is plagued by hallucinations again, apparently of a time some 80 years ago. The music was composed by a man named Joran Belar living in that time. When Dax goes into neural shock, Dr. Renhol rules that the Dax symbiont has to be removed at the host's expense if their condition doesn't improve. In an attempt to find out more to save Jadzia, Sisko and Bashir contact Yolad, Joran's brother. The official story is that Joran had allegedly been rejected by the Symbiosis Commission and then murdered the doctor who made that decision. But Yolad eventually discloses that his brother was indeed joined for no less than six months - a fact that has been purged from all records. The symbiont was Dax, and Dax's memory was erased likewise. The two officers also learn that not just a tiny percentage but half of all Trills are fit to be joined. Having obtained information that could put the Commission into a precarious position, they can convince Dr. Renhol to abstain from the removal of the Dax symbiont and to save Jadzia by re-integrating the suppressed memories.

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